Aaron Krochmal

Associate Professor of Biology
  • B.S., Biology, Union College (NY), 1996
  • M.S., Biology, New York University, 1998
  • Ph.D., Ecology and Systematics, Indiana State University, 2003
  • General Biology I and II
  • Comparative Vertebrate Anatomy
  • Comparative Animal Physiology
  • General Endocrinology
  • Research Techniques in Herpetology

Research Interests

I am a broadly-trained, integrative organismal biologist interested in the behavior, physiology, and ecology of reptiles. Specifically, I combine these disciplines to investigate how reptiles perceive, interact with, and navigate their environments. 


Selected Recent and Forthcoming Publications

(* - undergraduate coauthor) 

Roth, T.C., and A.R. Krochmal. 2018. Where the wild things are: a naturalistic approach to studying cognition in Chelonians. Box for: Wilkinson, A., and E. Glass. Cold-blooded cognition: How to get a tortoise out of its shell. In: Animal Cognition, F. Amici and N. Bueno eds., Cambridge University Press. In press.

Krochmal, A. R., Roth, T. C. and O’Malley, H*. 2018.  An empirical test of the role of learning in translocation. Anim Conserv.  21 (1), 36–44.

Krochmal, A.R. and Roth, T.C. 2017. From Comfort to Confidence: Modeling Science as a Process of Risk-Taking in the Classroom. In Risk-taking in higher education: The importance of negotiating intellectual challenge in the college classroom. Kelty, R. and Bunten, B, eds. Lanham, MD: Rowman & Littlefield.

Roth, T. C., Krochmal, A. R., Gerwig, IV, W. B*., Rush, S*., Simmons, N. T*., Sullivan, J. D*., and Wachter, K*. 2016. Using Pharmacological Manipulation and High-precision Radio Telemetry to Study Spatial Cognition in Free-ranging Animals. J. Vis. Exp. (117), e54790, doi:10.3791/54790     

Roth, T.C. and A.R. Krochmal. 2016. Pharmacological Evidence is Consistent with a Prominent Role of Spatial Memory in Complex Navigation. Proceedings of the Royal Society of London, B. 383 (1824)

Krochmal, A.R., T.C. Roth, S. Rush* and K. Wachter*. 2015. Outsmarting rapid environmental change: the role of cognition in navigation. Communicative and Integrative Biology. 8:6 e1052922 DOI:

Roth, Timothy C., and A R. Krochmal. Cognition-centered conservation as a means of advancing integrative animal behavior. Current Opinion in Behavioral Sciences 6 (2015): 1-6 DOI:

Roth, T.C., A.R. Krochmal, and Z. Nemeth. 2015. Thinking about change: an integrative approach for examining cognition in a changing world. Integrative and Comparative Biology. 55: 347-353 DOI:

Roth, T.C., and A.R. Krochmal. 2015The role of age-specific learning and experience for turtles navigating a changing landscape. Current Biology 25:333-337. DOI:; Cover article and cover photo. Featured in Berkeley Science Review.