Aaron Krochmal

Associate Professor of Biology
  • B.S., Biology, Union College (NY), 1996
  • M.S., Biology, New York University, 1998
  • Ph.D., Ecology and Systematics, Indiana State University, 2003
  • General Biology I and II
  • Comparative Vertebrate Anatomy
  • Comparative Animal Physiology
  • General Endocrinology
  • Research Techniques in Herpetology

Research Interests

I am a broadly-trained, integrative organismal biologist interested in the behavior, physiology, and ecology of reptiles. Specifically, I combine these disciplines to investigate how reptiles perceive, interact with, and navigate their environments. 


Selected Recent Popular Press Coverage

Animal learning, memory, and migration, on National Public Radio’s The Academic Minute

Agribusiness goes hand-in-hand with animal conservation in New Scientist

Students learn about how animals learn in The Chronicle of Higher Education (scroll down)


Selected Recent and Forthcoming Publications

(* - undergraduate coauthor) 

Roth, T.C., and A.R. Krochmal. 2018. Where the wild things are: a naturalistic approach to studying cognition in Chelonians. Box for: Wilkinson, A., and E. Glass. Cold-blooded cognition: How to get a tortoise out of its shell. In: Animal Cognition, F. Amici and N. Bueno eds., Cambridge University Press. In press.

Krochmal, A. R., Roth, T. C. and O’Malley, H*. 2018.  An empirical test of the role of learning in translocation. Anim Conserv.  21 (1), 36–44.

Krochmal, A.R. and Roth, T.C. 2017. From Comfort to Confidence: Modeling Science as a Process of Risk-Taking in the Classroom. In Risk-taking in higher education: The importance of negotiating intellectual challenge in the college classroom. Kelty, R. and Bunten, B, eds. Lanham, MD: Rowman & Littlefield.

Roth, T. C., Krochmal, A. R., Gerwig, IV, W. B*., Rush, S*., Simmons, N. T*., Sullivan, J. D*., and Wachter, K*. 2016. Using Pharmacological Manipulation and High-precision Radio Telemetry to Study Spatial Cognition in Free-ranging Animals. J. Vis. Exp. (117), e54790, doi:10.3791/54790     

Roth, T.C. and A.R. Krochmal. 2016. Pharmacological Evidence is Consistent with a Prominent Role of Spatial Memory in Complex Navigation. Proceedings of the Royal Society of London, B. 383 (1824) Featured cover article

Krochmal, A.R., T.C. Roth, S. Rush* and K. Wachter*. 2015. Outsmarting rapid environmental change: the role of cognition in navigation. Communicative and Integrative Biology. 8:6 e1052922 DOI:

Roth, Timothy C., and A R. Krochmal. Cognition-centered conservation as a means of advancing integrative animal behavior. Current Opinion in Behavioral Sciences 6 (2015): 1-6 DOI:

Roth, T.C., A.R. Krochmal, and Z. Nemeth. 2015. Thinking about change: an integrative approach for examining cognition in a changing world. Integrative and Comparative Biology. 55: 347-353 DOI:

Roth, T.C., and A.R. Krochmal. 2015The role of age-specific learning and experience for turtles navigating a changing landscape. Current Biology 25:333-337. DOI:; Cover article and cover photo. Featured in Berkeley Science Review.